Scale and mealybug diagnostic workshop resources

Scale and Mealybug Diagnostic Workshop Resources

Resources from the Scale and Mealybug Diagnostic Workshop are listed below.

Due to its size, the document ‘Mealybugs of Central and South America’, by Williams and de Willink 1992, is been divided into the following sections:

Due to its size, the document ‘Scale and Mealybugs of California Part 3: The armored scales (Homoptera: Coccoidea: Diaspididae)’, by Gill 1997, has been divided into the following sections:

Coccoidea diagnostic workshop references


Ben-Dov 2011, An updated checklist of the scale insects (Hemiptera: Coccoidea) of the Margarodidae sensu lato group

Bhatti 1990, A New Monophlebine Genus (Homoptera: Coccoidea: Margarodidae: Monophlebinae) on Melaleuca L. in Australia

Gullan and Cook 2007, Phylogeny and higher classification of the scale insects (Hemiptera: Sternorrhyncha: Coccoidea)

Hodgson 2014, Phenacoleachia, Steingelia, Pityococcus and Puto – Neococcoids or Archaeococcoids? An Intuitive Phylogenetic Discussion Based on Adult Male Characters

Hodgson and Gounari 2006, Morphology of Marchalina hellenica (Gennadius) (Hemiptera: Coccoidea: Marchalinidae) from Greece, with a discussion on the identity of M. caucasica Hadzibeyli from the Caucasus

Lapolla et al 2008, A new ortheziid (Hemiptera: Coccoidea) from Australia associated with Acropyga myops Forel (Hymenoptera: Formicidae) and a key to Australian Ortheziidae

Morales 1991, Fauna of NZ no 21 Magarodidae (Insecta: Hemiptera)

Morrison 1928, A classification of higher groups and genera of the coccid family Margarodidae

Unruh and Gullan 2008, Identification guide to species in the scale insect tribe Iceryini, (Coccoidea: Monophlebidae)

Unruh and Gullan 2008, Molecular data reveal convergent reproductive strategies in iceryine scale insects (Hemiptera: Coccoidea: Monophlebidae), allowing the re interpretation of morphology and a revised generic classification

Williams and Gullan 2008, A revision of the Neotropical scale insect genus Protortonia Townsend (Hemiptera: Coccoidea: Monophlebidae: Llaveiini)

Hard scale

Dooley, Key to the genera of the Aspidiotini

Dooley, Key to the Diaspidini genera

Dooley, Key to the Parlatoriini genera of the Diaspididae

Ferris and Prabhaker 1947, The genus Pinnaspis cockerell (Homoptera: Coccoidea: Diaspididae)

McKenzie 1945, A revision of the Parlatoria and closely allied genera (Homoptera: Coccoidea: Diaspididae)

Smith-Pardo et al 2012, A review of the genus Chrysomphalus Ashmead (Hemiptera: Coccoidea: Diaspididae) with descriptions of a new species and a new, related genus

Watson 2015, Unaspis lansivora sp. n. (Hemiptera: Diaspididae), a new pest of Lansium domesticum (Meliaceae), and a key to Unaspis species


Cox 1983, An experimental study of morphological variation in mealybugs (Homoptera: Coccoidea: Pseudococcidae)

Cox 1989, The mealybug genus Planococcus (Homoptera: Pseudococcidae)

Hodgson et al 2008, Phenacoccus solenopsis Tinsley (Sternorrhyncha: Coccoidea: Pseudococcidae), an invasive mealybug damaging cotton in Pakistan and India, with a discussion on seasonal morphological variation

Kayden and Gullan 2012, A taxonomic revision of the mealybug genus Ferrisia Fullaway (Hemiptera: Pseudococcidae), with descriptions of eight new species and a new genus

Gimpel and Miller 1996, Systematic analysis of the mealybugs in the Pseudococcus maritimus complex (Homoptera: Pseudococcidae)

Gullan 2000, Identification of the immature instars of mealybugs (Hemiptera: Pseudococcidae) found on citrus in Australia

Williams 1989, The mealybug genus Rastrococcus Ferris (Hemiptera: Pseudococcidae)

Williams 2005, An account of the mealybug genus Paraputo Laing (Hemiptera: Coccoidea: Pseudococcidae) in the Pacific region

Other Neococcids

Chamberlain 1923, A systematic monograph of the Tachardiinae or lac insects (Coccidae)

Deitz 1979, Two new species of Colobopyga (Homoptera: Halimococcidae) from the Australian region

Hardy and Gullan 2007, A new genus and four new species of felt scales on Eucalyptus (Hemiptera: Coccoidea: Eriococcidae) in south-eastern Australia

Hodgson and Hardy 2013, The phylogeny of the superfamily Coccoidea (Hemiptera: Sternorrhyncha) based on the morphology of extant and extinct macropterous males

Hodgson and Miller 2002, A new subfamily, two new genera and three new species of Aclerdidae (Hemiptera: Coccoidea ) from southern Africa, with a phylogenetic analysis of relationships

Hodgson and Lagowska 2011, New scale insect (Hemiptera: Sternorrhyncha: Coccoidea) records from Fiji: three new species, records of several new invasive species and an updated checklist of Coccoidea

Hodgson and Williams 2016, A revision of the family Cerococcidae Balachowsky (Hemiptera: Sternorrhyncha, Coccomorpha) with particular reference to species from the Afrotropical, western Palaearctic and western Oriental Regions, with the revival of Antecerococcus Green and description of a new genus and fifteen new species, and with ten new synonomies

Kondo and Gullan 2007, Taxonomic review of the lac insect genus Paratachardina Balachowsky (Hemiptera: Coccoidea: Kerriidae), with a revised key to genera of Kerriidae and description of two new species

Kozar and Foldi 2004, Description of new genera and species in the tribe Rhizoecin (Homoptera, Coccoidea, Pseudococcidae)

Muniappan et al 2009, New Records of Invasive Insects (Hemiptera: Sternorrhyncha) in Southeast Asia and West Africa

Pellizzari and Germain 2010, Scales (Hemiptera, Superfamily Coccoidea) Chapter 9.3

Stumpf and Lambdin 1999, Taxonomic Status of Bambusaspis miliaris, B. robusta, and B. pseudomiliaris (Hemiptera: Coccoidea: Asterolecaniidae)

Soft scale

Evans, Soft Scales of Southeast Asia (Hemiptera: Coccidae)

Ben-Dov 1979, A taxonomic study of the soft-scale genus Kilifia (Coccidae)

Fetyko and Kozar 2012, Records of Ceroplastes Gray 1828 in Europe, with an identification key to species in the Palaearctic Region

Grimshaw and Donaldson 2007, New records of mango shield scale Milviscutulus mangiferae (Green) (Hemiptera: Coccidae) and Brevennia rehi (Lindinger) (Hemiptera: Pseudococcidae) in north Queensland

Lin et al 2017, A newly recognised species that has been confused with the global polyphagous pest scale insect, Coccus hesperidum Linnaeus (Hemiptera: Coccomorpha: Coccidae)

Qin and Gullan 1992, A revision of the Australian pulvinariine soft scales (Insecta: Hemiptera: Coccidae)

Qin and Gullan 1994, Taxonomy of the wax scales (Hemiptera: Coccidae: Ceroplastinae) in Australia

Rakimov et al 2013, Soft scale insects (Hemiptera: Coccoidea: Coccidae) on grapevines in Australia

Wang and Feng 2012, A new species of Coccus (Hemiptera: Coccoidea: Coccidae) from China

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